This is a close up of a llama eye. The strange "ruffles" are called 'iridic granules' (corpora nigra) and they are used to to shade the eye from bright sunlight. In bright light these iridic granules can actually interlock to completely cover the centre of the pupil. This leaves just two holes open on either end of the pupil, reducing the amount of light that can enter the eye. These odd looking structures are also found in horses, cattle and sheep.